Study of Human Anatomy, Otago University, Dunedin/New Zealand, PhD (1999), postdoctoral fellow at National Institute of Mental Health, Bethesda/USA (1999-2003) and at Max Planck Institute for Medical Research, Heidelberg (2003-2006), Group Leader at Max Planck Institute for Biological Cybernetics, Tübingen (2006-2013), Scientific member of the Max Planck Society, Director at the MPI for Neurobiology of Behaviour (previously, caesar).